Note: For many non-migratory species, occurrences are roughly equivalent to populations.
Estimated Number of Occurrences: > 300
Comments: It occurs in creeks and the Savannah River on property of the Savannah River Power Plant, Aiken, South Carolina (Wood, 1982). In Georgia, this species is found in the eastern coastal plain (Dillon et al., 2006). It is common in coastal drainages in North Carolina (Dillon et al., 2006) and South Carolina (Dillon and Stewart, 2003). In Alabama, it is common and presumably occurs throughout the state (Mirarchi, 2004). It is widely distributed in Missouri (Audrain, Benton, Cass, Clark, Crawford, Holt, Howard, Iron, Johnson, Lincoln, Putnam, Ralls, Ripley, St. Charles, Stoddard, Wayne Cos.) (Wu et al., 1997). In Kentucky, it has been documented in the Tradewater (Flat Creek) (Branson and Batch, 1983) and Kentucky (farm pond in Madison Co.) (Branson and Batch, 1981) drainages. Blair and Sickel (1986) documented it in 10 of 44 sites surveyed in Land Between the Lakes (national recreation area between Cumberland River (Lake Berkeley) and Tennessee River (Kentucky Lake)) in Kentucky and Tennessee (both). Branson and Batch (1987) documented it in Kentucky in the Cumberland drainage (Bert T. Combs Pond, Bell Co.). In Indiana, Pyron et al. (2008) found it in 9 of 123 sites surveyed and 9 historical sites primarily north of the Wabash drainage compared to every drainage historically (Goodrich and van der Schalie, 1944). Jokinen (2005) found it at Indiana Sand Dunes National Lakeshore in Indiana. Pearce and Evans (2008) documented it in Plummers Island (Middle Potomac River system), Maryland. In Pennsylvania, it occurs in the Delaware and Ohio basins (Evans and Ray, 2010). In Alberta it is found throughout the province (Lepitzki, 2001). Lysne and Pierce (2009) found it rare in Crystal Creek-Spring Creek, Blaine Co., Idaho. Lysne and Clark (2009) found it in the Bruneau River (survey area from Snake River confluence upstream to Hot Creek- 41 km) in Idaho. It occurs in Mexico in Morelos and Sonora (Thompson, 2008).