Find content specifically related to fishes of the Puget Sound and Salish Sea ecosystems. For checklists and descriptive accounts of individual species, visit our species library. The Encyclopedia of Puget Sound will also be creating additional pages and sections related to salmon recovery in the region. For a general overview of salmonids, visit the Encyclopedia's Puget Sound Science Review.
Chemicals, disease and other stressors can increase a salmon's chance of being eaten or reduce its ability to catch food. We wrap up our series on the Salish Sea Marine Survival Project with a look at some of the lesser-known, but still significant factors contributing to salmon declines in the Salish Sea.
Researchers are analyzing the harmful effects of creosote-treated wood pilings on Pacific herring and shellfish in Puget Sound. Studies show that piling removal projects can ease the impacts, but only if carefully done.
A recent influx of anchovies into Puget Sound may have saved some steelhead from predators, but researchers seek more evidence to prove the connection. Our series on the Salish Sea Marine Survival Project continues with a look at these and other potential impacts from predators on the region's salmon and steelhead.
Getting bigger faster can help save juvenile Chinook salmon from a gauntlet of hungry predators ranging from birds and marine mammals to larger fish. We continue our series on the Salish Sea Marine Survival Project with a look at what helps salmon grow and prepare for life in the open ocean.
An intensive research program in the U.S. and Canada is studying why so few salmon in the Salish Sea are returning home to spawn. They are uncovering a complex web of problems involving predators, prey and other factors that put salmon at risk as they migrate to the ocean. We begin a four-part series on the Salish Sea Marine Survival Project, including new findings presented at the 2018 Salish Sea Ecosystem Conference last spring in Seattle.
A high-profile salmon escape led to a ban on salmon farms in Washington earlier this year. But just across the border, scientists say salmon farms in British Columbia expose migrating fish from Puget Sound to potential maladies like parasites, bacteria and dangerous viruses. They say simply getting rid of salmon farms in Washington does not put the potential impacts to rest.
Pacific herring exposed to stormwater in Puget Sound show some of the same effects as fish exposed to major oil spills. Symptoms include heart and developmental problems.
As wildlife managers work to recover Puget Sound’s diminished Chinook population, a proposed white paper is expected to review the impacts of some of the salmon's chief predators. The study would include a section on potential management of seals and sea lions, prompting open discussion of a long taboo subject: Could officials seek to revise the Marine Mammal Protection Act — or even conduct lethal or non-lethal removal of seals and sea lions in some cases? Such actions are hypothetical, but we look at some of the ongoing discussions around the issue as prompted by a new resolution from the Puget Sound Leadership Council.
The Puget Sound Ecosystem Monitoring Program has released its sixth annual Marine Waters Overview. The report provides an assessment of marine conditions for the year 2016 and includes updates on water quality as well as status reports for select plankton, seabirds and fish.
Increased consumption of Chinook salmon by seals and sea lions in the Salish Sea “could be masking the success of coastwide salmon recovery efforts,” according to a new study published in the journal Scientific Reports. Endangered resident orcas are said to be declining in part due to a lack of available Chinook, the orcas' preferred prey.
The 2017 State of the Sound is the Puget Sound Partnership’s fifth biennial report to the Legislature on progress toward the recovery of Puget Sound by 2020. The document reports on both the status of the Partnership's recovery efforts and the status of a suite of ecosystem indicators.
Formaldehyde is often used to control parasites on hatchery salmon and trout. The U.S. Environmental Protection Agency and the Washington State Department of Ecology conducted a joint study of formaldehyde concentrations in effluent from hatcheries in the Pacific Northwest.
A new study shows a surprising decline in some toxic chemicals in Puget Sound fish, while levels of PCBs increased in some cases. Scientists say the study shows that banning toxic chemicals can work, but old contaminants remain a challenge as they continue to wash into Puget Sound.
Toxic chemicals have been showing up in Puget Sound fish for more than a century, but consistent testing over the past 30 years has helped to reveal some unusual patterns of pollution.
In recent decades, hundreds of millions of dollars have been spent to restore habitat for Puget Sound salmon. In this article, we look at how scientists are gauging their progress. Are environmental conditions improving or getting worse? The answer may depend on where you look and who you ask.
A biennial report produced by the Governor's Salmon Recovery Office provides stories and data about salmon, habitat, and salmon recovery in Washington, including Puget Sound.
The marine habitat of Puget Sound can be divided up into nearshore, benthic (associated with the sea floor), and pelagic (open water) habitats. This article focuses on the pelagic habitat within the Puget Sound. This article was prepared as part of the 2015 Puget Sound Fact Book produced by the University of Washington Puget Sound Institute.
A 2017 report from the Puget Sound Ecosystem Monitoring Program presents an overview of selected recent monitoring and research activities focused on toxic contaminants in the Salish Sea.
The Puget Sound Ecosystem Monitoring Program (PSEMP) is an independent program established by state and federal statute to monitor environmental conditions in Puget Sound.
A new approach to flood control is taking hold across Puget Sound. Rivers, scientists say, can be contained by setting them free. Conservationists hope this is good news for salmon recovery.
As threatened Chinook populations in Puget Sound continue to lose ground, the state is looking to new strategies to reverse the trend. In the Skagit watershed, the scientists — and the fish — are among those leading the way.
A new study shows that increased populations of seals and sea lions are eating far more of Puget Sound’s threatened chinook than previously known, potentially hampering recovery efforts for both salmon and endangered killer whales.
Sand lance in parts of British Columbia are ingesting small pieces of plastic that may be passed through the food web.
Some of the most important fish in the Salish Sea food web are also the most mysterious. Researchers have only begun to understand how many there are, where they go, and how we can preserve their populations for the future. A University of Washington researcher describes how scientists are looking into the problem.
New, smaller acoustic tags will allow scientists to track steelhead migrations in Puget Sound in ways that were once impossible. Will they provide answers to the mysterious decline of these now-threatened fish?
The Salish Sea Marine Survival Project has mobilized dozens of organizations in the U.S. and Canada to find an answer to one of the region's greatest mysteries. What is killing so many young salmon before they can return home to spawn? A series of talks at the 2016 Salish Sea Ecosystem Conference brought together some of the latest research.
Many of Puget Sound's Chinook salmon spend their entire lives in local waters and don't migrate to the open ocean. These fish tend to collect more contaminants in their bodies because of the sound's relatively high levels of pollution.
A 2015 report from Snohomish County, King County and the Tulalip Tribes outlines protection strategies for salmon and salmon habitat within the Snohomish Basin.
Researchers are proposing a shift in thinking about how some of the region’s most damaging pollutants enter Puget Sound species like herring, salmon and orcas.
The growing number of species of concern in the Salish Sea suggests ecosystem decay is outpacing recovery
The number of species of concern in the Salish Sea is growing at an average annual rate of 2.6%, according to a report published in the proceedings of the 2016 Salish Sea Ecosystem Conference in Vancouver, B.C.
The number of species of concern in the Salish Sea is growing at an average annual rate of 2.6%, according to a report published in the proceedings of the 2016 Salish Sea Ecosystem Conference in Vancouver, B.C.
Scientists are realizing that underwater noise in the Salish Sea affects a broad range of species, even plankton. Read a Q & A with the organizers of the session 'From plankton to whales: underwater noise and its impacts on marine life.'
Where shoreline bulkheads remain in place, the loss of spawning habitat used by surf smelt is likely to reach 80 percent.
Rising sea levels are expected to exacerbate habitat loss caused by bulkheads, according to studies in the San Juan Islands.
A 2016 paper in Environmental Pollution identifies dozens of pharmaceuticals and other compounds that are accumulating in Puget Sound fish such as salmon.
A 2016 paper in the journal Oecologia describes how individual herring populations in Puget Sound exhibit a portfolio effect, collectively influencing and stabilizing the region’s population as a whole.
Food webs are natural interconnections of food chains and depict what-eats-what in an ecological community. While Puget Sound represents a specific food web, the organisms that reside within that web often travel outside the region. In this way, one community's food web can be drastically affected by a change in a neighboring ecosystem.
Decaying organic matter plays an important role in marine ecosystems.
Chinook, coho and sockeye salmon, along with steelhead trout, live in the Lake Washington watershed and navigate a treacherous route through the Ballard Locks on their way to Puget Sound.
Lake Washington was heavily contaminated by untreated sewage until extensive pollution controls by the city of Seattle.
Impacts of anthropogenic noise on marine life: Publication patterns, new discoveries, and future directions in research and management
A 2015 review in Ocean & Coastal Management looks at trends in research related to anthropogenic noise and its affect on a wide variety of marine organisms, from whales and fish to invertebrates. The review includes case studies from the Salish Sea.
Pacific herring have long been considered an essential part of the Puget Sound food web. Now, studies are beginning to reveal how diseases in herring could be reverberating through the ecosystem, affecting creatures large and small. We continue our coverage of the ecological impacts of disease in Puget Sound with this look at the region's most well-known forage fish.
Chinook, coho and steelhead populations in Puget Sound have declined dramatically over the past 30 years. In some cases, counts of fish returning to the rivers are just a tenth what they were in the 1980s. While many possible causes of this decline are under consideration, some researchers are focusing on the combined effects of predators and disease. This article continues our coverage of the ecological impacts of disease in Puget Sound.
Researchers updating a 1980 fish catalog have found evidence of 37 additional fish species in the Salish Sea. This information, accompanied by hundreds of detailed illustrations, is seeding a new reference book expected to gain wide use among scientists, anglers and conservationists.
The 2015 Puget Sound Fact Book brings together statistics and other information about the health and makeup of the Puget Sound ecosystem. Areas of focus include climate change, geography, water quality, habitats, human dimensions and regional species. The fact book was prepared for the Encyclopedia of Puget Sound with funding from the Environmental Protection Agency and the Puget Sound Partnership.
A 2015 NOAA report creates an updated and comprehensive list of the fishes of the Salish Sea.
A report from NOAA and the Puget Sound Ecosystem Monitoring Program provides an overview of 2014 marine water quality and conditions in Puget Sound from comprehensive monitoring and observing programs.
This report documents how Washingtonians have responded to the challenges of protecting and restoring salmon and steelhead to healthy status. It also serves as a tool to summarize achievements, track salmon recovery progress through common indicators, and identify data gaps that need to be filled.
Review of the marine environment and biota of Strait of Georgia, Puget Sound and Juan de Fuca Strait
Proceedings of the BC/Washington Symposium on the Marine Environment, January 13 and 14, 1994
Conservation and ecology of marine forage fishes— Proceedings of a research symposium, September 2012
The symposium was held on September 12–14, 2012, at the University of Washington, Friday Harbor Laboratories campus. Sixty scientists, graduate students, and fisheries policy experts convened; showcasing ongoing research, conservation, and management efforts targeting forage fish from regional and national perspectives.
Salmon recovery demands both dedication among people with different interests, and sustained resources. This biennial report tells the story of the progress made to date and the challenges ahead.
The Puget Sound Update is a technical report that integrates results of PSAMP and other scientific activities in Puget Sound focused on marine life and nearshore habitat, marine and freshwater quality, and toxic contamination.
A 2015 paper in the journal Ecological Economics evaluated “personal use” and subsistence use of seafood among commercial operators in Washington and California, as well as the extent, range, and species diversity of noncommercial wild ocean seafood subsistence harvests.
Monitoring for adaptive management: status and trends of aquatic and riparian habitats in the Lake Washington/Cedar/Sammamish watershed (WRIA 8)
King County conducted physical and biological monitoring between 2010 and 2013 in the Lake Washington/Cedar/Sammamish (WRIA 8) watershed using common survey protocols and a probabilistic survey design. Hydrologic monitoring was also conducted at several locations to supplement physical and biological monitoring.
Scientists say low marine survival rates threaten Puget Sound coho salmon populations. A 2015 article in the journal Marine and Coastal Fisheries reports that wild cohos in the Salish Sea had higher smolt survival rates over a 30 year period than hatchery coho salmon. Smolt survival in the Strait of Georgia during that time declined faster than it did in Puget Sound.
Forty years of change in forage fish and jellyfish abundance across greater Puget Sound, Washington (USA): anthropogenic and climate associations
A 2015 paper in the Marine Ecology Press Series reports a trend toward more jellyfish and less of some forage fish species in Puget Sound. The paper analyzes more than 40 years of state data, and assesses potential human causes for the shift.
A 2015 paper in Oikos Journal examines the impacts of great blue heron predation on species diversity in eelgrass meadows in British Columbia.
A 2015 paper in the journal Marine Policy examines surveys of Puget Sound anglers to provide baseline information related to rockfish conservation.
A 2014 report by the North Cascadia Adaptation Partnership identifies climate change issues relevant to resource management in the North Cascades, and recommends solutions that will facilitate the transition of the diverse ecosystems of this region into a warmer climate.
How does one of the West's busiest airports deal with extreme stormwater, and what does that mean for water quality standards in the rest of the state?
This 2006 technical report for the Puget Sound Nearshore Partnership describes how shellfish have high ecological, economical, cultural, recreational value, however human activity is threatening their existence by altering their native habitat with changes in land use, shoreline modifications, stormwater, sewage and industrial discharge.
Scientists are rethinking floodplain management in Puget Sound. Can we have our farms and salmon too?
Puget Sound herring reproduction is not limited by the amount of suitable spawning vegetation, according to a November 2014 paper in the journal Marine Ecology. The article points to terrestrial or marine variables as likely determinants of egg loss.
The decaying seawall along Seattle’s waterfront is providing scientists with an opportunity to improve long-lost habitat for migrating salmon. It could also show the way for habitat enhancements to crumbling infrastructure worldwide. One University of Washington researcher describes the project.
A 2014 paper decribes how monitoring the energy density of key Pacific salmon species could affect the recovery of northern and southern killer whales through fisheries management.
Monitoring and adaptive management of the Nisqually Delta after tidal marsh restoration: Restoring ecosystem function for salmon
This 2009 report by the Nisqually Tribe establishes key measures of restoration development, habitat processes, and Chinook salmon response for the largest delta restoration project in the Pacific Northwest.
Birds that dive for fish while wintering in the Salish Sea are more likely to be in decline than nondiving birds with less specialized diets, according to a 2014 study led by the School of Veterinary Medicine at the University of California, Davis. The study, published in the journal Conservation Biology proposes that long-term changes in the availability of forage fish are pushing the declines.
New research presented at the 2014 Salish Sea Ecosystem Conference shows that some of the greatest dangers to Puget Sound marine life come from our common, everyday activities. These pervasive sources of pollution are so woven into our lives that they are almost invisible to us, but it’s becoming impossible to ignore their effects.
The growth and survival of young salmon in streams, river deltas and floodplains are seen as crucial pieces of the salmon recovery puzzle. In part two of this two-part series, researchers at the Salish Sea Ecosystem Conference in Seattle say the complexities of the salmon life cycle require new coordination among scientists.
Scientists say Puget Sound’s salmon are dying young and point to low growth rates in the marine environment as a possible cause. In part one of this two-part series, scientists consider threats facing young salmon in the open waters of Puget Sound.
Scientists say eelgrass, an unassuming flowering plant found just off shore in Puget Sound, is vital to the health of the ecosystem. They also say the plant is declining. New and increasingly urgent efforts to restore it brought a group of researchers to the 2014 Salish Sea Ecosystem Conference.
Spatial and Temporal Variation in River Otter (Lontra canadensis) Diet and Predation on Rockfish (Genus Sebastes) in the San Juan Islands, Washington
A 2014 paper in the journal Aquatic Mammals examines coastal river otter predation on rockfish at three islands in the Salish Sea.
Approximately every two years, the SeaDoc Society prepares a list of species of concern within the Salish Sea ecosystem. The following paper found 119 species at risk and was presented as part of the proceedings of the 2014 Salish Sea Ecosystem Conference, April 30 – May 2, 2014, Seattle, Washington.
A 2014 report describes a study of socio-cultural values associated with blueback salmon in the Quinault Indian Nation. The blueback salmon (Oncorhynchus nerka) is a unique strain of sockeye that returns primarily to the Quinault river system.
Lead Entities are local organizations in Puget Sound that develop salmon recovery strategies and priorities for the region on a watershed-based scale.
Forage fish represent a critical link in the Puget Sound food web and help to sustain key species like salmon, marine mammals and sea birds. But the region’s forage fish may be vulnerable on a variety of fronts, according to a new study panel report from the University of Washington Puget Sound Institute. Download the panel's summary and proposed research plan.
An October 2013 report released by the Puget Sound Ecosystem Monitoring Program inventories and assesses monitoring activities of Puget Sound's ESA-listed salmon and steelhead stocks and habitats.
The Encyclopedia of Puget Sound spoke with Seattle Times reporter Lynda Mapes about the exhibit Elwha: A River Reborn, which opened at the University of Washington Burke Museum on November 23rd. The exhibit is based on the book of the same title by Mapes and photographer Steve Ringman, and tells the story of the largest dam removal in U.S. history.
This article was originally published by the Washington Department of Fish and Wildlife as part of its annual report Threatened and Endangered Wildlife in Washington.
A camera on board a remotely operated vehicle scans the floor of Puget Sound capturing digital video of underwater marine life. Selected clips of Plumose sea anemones, Pacific halibut, Pacific cod, Sea stars, and North Pacific spiny dogfish are now available for public viewing.
Download presentations from the Study Panel on Ecosystem-based Management of Forage Fish held August 25, 2013 at the University of Washington’s Friday Harbor Lab, San Juan Island.
Paper: A floating bridge disrupts seaward migration and increases mortality of Steelhead smolts in Hood Canal, Washington State
A new study provides strong evidence of substantial migration interference and increased mortality risk associated with the Hood Canal Bridge for aquatic animals, and may partially explain low early marine survival rates observed in Hood Canal steelhead populations.
Remotely Operated Vehicles (ROVs) and Autonomous Underwater Vehicles (AUVs) are underwater robotic vehicles used for a variety of ocean surveys and operations. Both are used for deep-sea observation, mapping of underwater environments, and surveys of biodiversity and water quality trends. While ROVs are tethered to the user by a cord called the umbilical, which provides power as well as control and video signals, AUVs are programmed for a specific course and then set loose, operating without a tether.
Proposed designation of critical habitat for the distinct population segments of Yelloweye Rockfish, Canary Rockfish, and Bocaccio
The National Marine Fisheries Service has released a Draft Biological Report proposing designation of critical habitat for yelloweye rockfish, canary rockfish, and bocaccio in the Salish Sea. Download the full report and supporting data.
The Puget Sound Marine Waters 2012 Overview from the Puget Sound Ecosystem Monitoring Program synthesizes conditions measured in 2012 and has been expanded to include observations on seabirds that rely on marine waters. Read an excerpt below, or download the full report.
The audio files below are excerpts from a May 2013 interview with Donald Malins, former Director of the Environmental Conservation Division of NOAA Fisheries. Research by Malins and his colleagues in the 1970s and mid-1980s revealed high levels of industrial toxics in sediment-dwelling fish in Puget Sound, leading to the creation of Superfund sites in the Duwamish Estuary and Commencement Bay. Read a full profile of Donald Malins. The interview was conducted by Richard Strickland and Randy Shuman in cooperation with the Encyclopedia of Puget Sound as part of the Puget Sound Voices series. Additional assistance was provided by Jake Strickland.
In the 1970s and 1980s, research from a division of NOAA's Montlake Lab suddenly and irreversibly changed the way scientists and the public viewed the health of Puget Sound. Their discoveries of industrial toxics in the region's sediment-dwelling fish led to the creation of two Superfund sites, and new approaches to ecosystem management across the Sound. The man at the forefront of this research was Dr. Donald Malins, featured here as part of the Puget Sound Voices series.
Paper: Shifts in the estuarine demersal fish community after a fishery closure in Puget Sound, Washington
This paper looks at 21 years of data on estuarine demersal fish in Puget Sound, assessing changes in population after the closure of bottom trawl fisheries.
Could recent declines in Puget Sound herring be linked to decreases in native eelgrass? Biologist Tessa Francis reports on a new study that may provide insight into the health of one of the region's most iconic forage fish.
Report: Economic analysis of the non-treaty commercial and recreational fisheries in Washington State
This report, published in 2008 by the Washington Department of Fish and Wildlife, summarizes the economic importance of Washington fisheries using data from 2006. The report's Executive Summary is reprinted below, followed by summaries of data specific to Puget Sound.
This document was prepared by the U.S. Geological Survey in cooperation with the U.S. Environmental Protection Agency and the National Park Service. Download the entire report, or read the Introduction below. Portions of this document were originally published in June 2013 and were updated in February 2014.
Seabird populations are declining worldwide. This paper looks at the impact of gillnets on bird populations.
A master's thesis prepared at Western Washington University discusses the impact of harbor seals on fish stocks in the San Juan Islands, where the seals are a year-round predator.
With funding from the EPA (EPA Interagency Agreement DW-13-923276-01), scientists at the Northwest Fisheries Science Center and the University of Washington used a field and quantitative modeling ‘source-transport-fate’ assessment approach to classify the vulnerability of shellfish growing areas to closures caused by watershed and marine-derived pathogens. Based on the historical prevalence of nutrient pollution, shellfish closures, and phytoplankton blooms in commercial and recreational shellfish growing area, the project focused on three nearshore sites--the Hamma Hamma (WRIA 16), Dosewallips (WRIA 16) and Samish (WRIA 3).
Extended abstract— Poisoning the body to nourish the soul: Prioritising health risks and impacts in a Native American community
This is an extended abstract of Poisoning the body to nourish the soul: Prioritising health risks and impacts in a Native American community by Jamie L. Donatuto, Terre A. Satterfield and Robin Gregory. The full article was published in Health, Risk & Society, Vol. 13, No. 2, April 2011, 103–127. The extended abstract was prepared for the Encyclopedia of Puget Sound by Jamie L. Donatuto.
A new Chinook monitoring framework is designed to build cooperation among managers and policymakers working across the Puget Sound watershed. The report, prepared by an independent team of scientists and released by NOAA, includes a regionally specific, common classification system for Chinook habitats and key ecological attributes.
Puget Sound Chinook Salmon recovery: a framework for the development of monitoring and adaptive management plans
The Puget Sound Recovery Implementation Technical Team has released a draft of a NOAA technical memorandum describing frameworks for adaptive management and monitoring of Chinook salmon in Puget Sound. Download the report.
This is the executive summary from a technical report produced for the Puget Sound Nearshore Partnership on Valued Ecosystem Components (VEC). The entire document is included as a PDF with this summary.
Canadian and U.S. governments differ on special status for bocaccio in the Salish Sea.
The Encyclopedia of Puget Sound species library now includes a list of species of concern in the Salish Sea watershed. The list was created by Joe Gaydos and Nicholas Brown of the SeaDoc Society, and was released as a paper presented as part of the Proceedings of the 2011 Salish Sea Ecosystem Conference in Vancouver, BC.
Methods and Quality of VSP Monitoring Of ESA Listed Puget Sound Salmon and Steelhead: With Identified Critical Gaps 2012
There are at least 28 species of rockfish in the Salish Sea, but their populations have declined in the past several decades. The proceedings from a 2011 rockfish recovery workshop in Seattle are now available.
A recent report by an independent science panel reviewed data on the effects of salmon fisheries on Southern Resident Killer Whale populations. The report was released on November 30, 2012 and was commissioned by NOAA Fisheries and Oceans Canada.
Fish in the family Salmonidae (salmon, trout, and charr) play potentially integral roles in the upland freshwater, nearshore and pelagic marine ecosystems and food webs of Puget Sound.
Approximately 28 species of rockfish are reported from Puget Sound, spanning a range of life-history types, habitats, and ecological niches.
Bentho-pelagic fish utilize both bottom habitats and shallower portions of the water column, often feeding in shallow water at night and moving to deeper water to form schools during the day.
The State of Our Watersheds Report is produced by the treaty tribes of western Washington, and seeks to present a comprehensive view of 20 watersheds in the Puget Sound region and the major issues that are impacting habitat.
The Puget Sound Marine Waters 2011 report is now available. The report was produced by the Puget Sound Ecosystem Monitoring Program and assesses the condition and quality of the waters of Puget Sound.
Forage fish occupy every marine and estuarine nearshore habitat in Washington, and much of the intertidal and shallow subtidal areas of the Puget Sound Basin are used by these species for spawning habitat.
Puget Sound has over 4,000 km (2,500 miles) of shorelines, ranging from rocky sea cliffs to coastal bluffs and river deltas. The exchange of water, sediment, and nutrients between the land and sea is fundamental to the formation and maintenance of an array of critical habitat types.
Puget Sound hosts more than 100 species of seabirds, 200 species of fish, 15 marine mammal species, hundreds of plant species, and thousands of invertebrate species. These species do not exist in isolation, but rather interact with each other in a variety of ways: they eat and are eaten by each other; they serve as vectors of disease or toxins; they are parasitic; and they compete with each other for food, habitat, and other resources.
A variety of animals, including invertebrates, fish, mammals, and birds, consume the suspension-feeders, filter-feeders, grazers, and detritivores that serve as a link between the primary producers and detrital pathways and the upper levels of the food web.
Fishes, birds, and mammals (including humans) serve as top-level carnivores in the Puget Sound ecosystem. With the exception of humans, these organisms have a diet that consists almost entirely of fish or other vertebrates.
Non-native species are those that do not naturally occur in an ecosystem. A non-native species is considered invasive when it is capable of aggressively establishing itself and causing environmental damage to an ecosystem. Plants, animals, and pathogens all can be invasive.