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Fishes, birds, and mammals (including humans) serve as top-level carnivores in the Puget Sound ecosystem. With the exception of humans, these organisms have a diet that consists almost entirely of fish or other vertebrates.

A variety of animals, including invertebrates, fish, mammals, and birds, consume the suspension-feeders, filter-feeders, grazers, and detritivores that serve as a link between the primary producers and detrital pathways and the upper levels of the food web.

Many consumer organisms in Puget Sound are both herbivores and detritivores. Zooplankton and benthic invertebrates that are scavengers, herbivores, or detritivores are considered jointly in this article. Some of these organisms can be predatory as well. Hundreds of invertebrates and fish species have a planktonic larval stage that eats plants and occupies the nearshore and offshore pelagic waters of Puget Sound.

Puget Sound hosts more than 100 species of seabirds, 200 species of fish, 15 marine mammal species, hundreds of plant species, and thousands of invertebrate species. These species do not exist in isolation, but rather interact with each other in a variety of ways: they eat and are eaten by each other; they serve as vectors of disease or toxins; they are parasitic; and they compete with each other for food, habitat, and other resources.

Ecosystem services are the “outputs” and experiences of ecosystems that benefit humans, and are generated by the structure and function of natural systems, often in combination with human activities. The Millennium Ecosystem Assessment, a global effort to catalog and assess ecosystem status and functions, offers a useful classification scheme.

The climate of Puget Sound is a product of the interaction between large-scale wind and weather patterns and the complex topography of the region. Seasonal changes in the movement of moisture-laden air that collides with the sudden barrier of the Olympic and Cascade mountains bring Puget Sound the record-breaking precipitation for which it is so famous. These circulation and topographic differences also lead to remarkable climate differences within Puget Sound itself, influencing the species and habitats that are found in the Sound.

Puget Sound has over 4,000 km (2,500 miles) of shorelines, ranging from rocky sea cliffs to coastal bluffs and river deltas. The exchange of water, sediment, and nutrients between the land and sea is fundamental to the formation and maintenance of an array of critical habitat types.