Note: For many non-migratory species, occurrences are roughly equivalent to populations.
Estimated Number of Occurrences: > 300
Comments: It occurs in creeks and the Savannah River on property of the Savannah River Power Plant, Aiken, South Carolina (Wood, 1982). In Georgia (Dillon et al., 2006), South Carolina (Dillon and Stewart, 2003), and North Carolina (Dillon et al., 2006; Dawley, 1965), it is known from the Blue Ridge and Eastern Piedmont regions but is generally absent from the Coastal Plain. Pearce and Evans (2008) documented it in Plummers Island (Middle Potomac River system), Maryland. In Pennsylvania, it occurs statewide and very common (Evans and Ray, 2010). Distribution in Alabama is poorly known but it is presumably found throughout the state (Mirarchi, 2004). In the Cuyahoga Valley National Park (between Cleveland and Akron, Ohio), this species was recently found in Cuyahoga River mainstem and at 7 tributaries (Smith et al., 2002). In Indiana, Pyron et al. (2008) found it in 18 of 123 sites surveyed compared to in Lake Knox and Henry Cos. historically (Goodrich and van der Schalie, 1944). It is fairly common in New York (Jokinen, 1992). It has been recorded throughout the Salt Fork of the Big Vermilion River, Illinois (Dexter, 1961). It has been found in Frog Bayou (Lower Arkansas drainage), Arkansas (Gordon, 1985). It occurs in the White River drainage, Arkansas, at the mouth in Washington Co. and in Independence Co. (Gordon, 1982). Minton et al. (2008) documented it in the Bayou Bartholomew drainage in Arkansas. In Missouri, it is only known from the Ozark Region in Crawford, Howell, Jasper, McDonald, Newton, Oregon, Ripley, and Wright Cos. (Wu et al., 1997) and the Spring River (Branson, 1966). In Kentucky it has been documented in the Salt River (Rolling Fork Salt River, Salt River) (Branson and Batch, 1983) and Licking (Licking River) River (Branson and Batch, 1981) drainages. Lysne and Pierce (2009) found it rare in Crystal Creek-Spring Creek, Blaine Co., Idaho. Lysne and Clark (2009) found it in the Bruneau River (survey area from Snake River confluence upstream to Hot Creek- 41 km) in Idaho.
